1. Oklahoma Department of Wildlife Conservation Oklahoma Fishery Research Laboratory 500 East Constellation Norman Oklahoma 73072 USA
2. U.S. Fish and Wildlife Service, Division of Biological Sciences 500 Gold Avenue Southwest Albuquerque New Mexico 87103‐1306 USA
3. U.S. Geological Survey Oklahoma Cooperative Fish and Wildlife Research Unit Department of Natural Resource Ecology and Management 007 Agriculture Hall, Oklahoma State University Stillwater Oklahoma 74078 USA