1. Aquatic Ecology Laboratory Department of Evolution Ecology, and Organismal BiologyThe Ohio State University1314 Kinnear Road Columbus Ohio 43212USA
2. Illinois Department of Natural Resources 11731 State Highway 37 Benton Illinois 62812USA
3. Aquatic Ecology Laboratory The Ohio State University1314 Kinnear Road Columbus Ohio 43212USA
4. U.S. Geological Survey Kansas Cooperative Fish and Wildlife Research Unit Division of Biology Kansas State University Manhattan Kansas 66506 USA
5. U.S. Geological Survey Vermont Cooperative Fish and Wildlife Research Unit Rubenstein School of Environment and Natural Resources University of Vermont Burlington Vermont 05405 USA