1. Ecologie Systématique Evolution Université Paris‐Sud Orsay Cedex France
2. CNRS, AgroParisTech Université Paris‐Saclay Orsay Cedex France
3. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences Beijing Normal University Beijing China
4. Department of Bioscience Aarhus University Rønde Denmark