1. Department of Earth, Ocean and Ecological Sciences University of Liverpool LiverpoolL69 3GPUK
2. Department of Zoology and Entomology University of Pretoria Pretoria0002South Africa
3. Environmental Change Institute School of Geography University of Oxford OxfordOX1 3QYUK
4. Department of Biological Sciences University of Cyprus Nicosia Cyprus
5. School of Animal, Plant and Environmental Sciences University of the Witwatersrand Wits South Africa