1. Department of Biology Amherst College Amherst Massachusetts
2. Department of Biology University of Virginia Charlottesville Virginia
3. Ecologie Systematique et Evolution, Univ. Paris‐Sud, CNRS, AgroParisTech Université Paris Saclay Orsay France
4. INRA ‐ UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier SupAgro) Montferrier‐sur‐Lez France