1. Research Institute for the Environment and Livelihoods; Charles Darwin University; Darwin Northern Territory 0909 Australia
2. PMB 44, Tropical Ecosystems Research Centre; CSIRO Land & Water; Winnellie Northern Territory 0822 Australia
3. Departamento de Botanica; Universidade Federal de Pernambuco; Av. Prof. Moraes Rego s/no Recife Pernambuco 50670-901 Brazil
4. School of Biological Sciences; University of Western Australia; Perth Western Australia 6009 Australia