1. Guangxi Key Laboratory of Marine Environmental Science, Guangxi Beibu Gulf Marine Research Center Guangxi Academy of Sciences Nanning Guangxi China
2. Laboratory of Marine Biology and Ecology Third Institute of Oceanography, Ministry of Natural Resources Xiamen China
3. College of Marine Sciences Beibu Gulf University Qinzhou Guangxi China
4. Clymene Enterprises Yerba Valley Way Lakeside California USA