1. Fish Ecology and Conservation Physiology Laboratory; Department of Biology and Institute of Environmental Science; Carleton University; 1125 Colonel By Drive Ottawa Ontario K1S 5B6 Canada
2. Ontario Ministry of Natural Resources and Forestry; 3301 Trout Lake Road North Bay Ontario P1B 8G4 Canada
3. Department of Environmental Conservation; University of Massachusetts Amherst; Amherst Massachusetts 01003 USA