1. Department of Pharmacology, Faculty of MedicineUniversity of Miyazaki Miyazaki Japan
2. Department of BiochemistryKyushu University Graduate School of Medical Sciences Fukuoka Japan
3. Laboratory for Animal Resource Development, RIKEN Center for Biosystems Dynamics Research Kobe Japan
4. Laboratory for Genetic EngineeringRIKEN Center for Biosystems Dynamics Research Kobe Japan