1. Key Laboratory of Sichuan Province for Fishes Conservation and Utilization in the Upper Reaches of the Yangtze River, and College of Life Science Neijiang Normal University Neijiang Sichuan 641000 People’s Republic of China
2. Neijiang City Academy of Agricultural Sciences Neijiang Sichuan 641000 People’s Republic of China
3. College of Life Science Neijiang Normal University Neijiang Sichuan 641000 People’s Republic of China
4. Engineering Research Center of Eco‐environment in Three Gorges Reservoir Region Ministry of Education China Three Gorges University Yichang Hubei 443002 People’s Republic of China