1. Department of Ecology, Evolution, and Marine Biology University of California Santa Barbara Santa Barbara California93101USA
2. National Center for Ecological Analysis and Synthesis 735 State Street Santa Barbara California93101USA
3. Conservation Biology Division Northwest Fisheries Science Center National Marine Fisheries Service National Oceanic & Atmospheric Administration Seattle Washington98112USA
4. School of Environmental and Forest Sciences University of Washington Box 355020 Seattle Washington98195USA
5. The Nature Conservancy 74 Wall Street Seattle Washington USA