1. School of Biology; University of St. Andrews; St. Andrews; KY16 8LB; UK
2. Institute of Biological; Environmental and Rural Sciences; Aberystwyth University; Aberystwyth; Ceredigion; SY23 3DA; UK
3. Department of Biology; University of Leicester; Leicester; LE1 7RH; UK
4. Department of Ecology and Vertebrate Zoology; University of Łódź; Łódź; Poland