1. Centre for Social Evolution; Department of Biology; University of Copenhagen; Universitetsparken 15 2100 Copenhagen Denmark
2. Smithsonian Tropical Research Institute; Apartado 0843-03092 Balboa Ancon Republic of Panama
3. CREAF; Cerdanyola del Vallès ES-08193 Catalunya Spain
4. Terrestrial Ecology Section; Department of Biology; University of Copenhagen; Universitetsparken 15 2100 Copenhagen Denmark