1. Department of Ecology and Evolutionary Biology University of California Santa Cruz Santa Cruz California 95064 USA
2. Elkhorn Slough National Estuarine Research Reserve Royal Oaks California 95076 USA
3. Department of Environmental Engineering Sciences Engineering School of Sustainable Infrastructure and Environment University of Florida Gainesville Florida 32611 USA
4. Division of Marine Science and Conservation Nicholas School of the Environment Duke University Beaufort North Carolina 28516 USA
5. Department of Biology Sonoma State University Rohnert Park California 94928 USA