1. Department of Biological Sciences; Graduate School of Science; Hiroshima University; Higashi-Hiroshima Hiroshima Japan
2. Institute for Amphibian Biology; Graduate School of Science; Hiroshima University; Higashi-Hiroshima Hiroshima Japan
3. Aitsu Marine Station; Center for Marine Environmental Studies; Kumamoto University; Kami-Amakusa Kumamoto Japan