1. Department of Biology; University of Waterloo; 200 University Avenue West Waterloo Ontario N2L 3G1 Canada
2. Fisheries and Oceans Canada; Great Lakes Laboratory for Fisheries and Aquatic Sciences; 1219 Queen Street East Sault Ste. Marie Ontario P6A 2E5 Canada
3. Fisheries and Oceans Canada; Ecosystem Science Directorate; 200 Kent Street Ottawa ON, K1A 0E6 Canada