1. Division of Mountain and Environmental Science, Interdisciplinary Graduate School of Science and Technology Shinshu University Matsumoto Japan
2. Division of Evolutionary Developmental Biology National Institute for Basic Biology Okazaki Japan
3. Department of Biology, Faculty of Science Shinshu University Matsumoto Japan
4. Hokkaido Aquatic Biology Eniwa Japan
5. Nagatsuta Yokohama Japan
6. Institute of Mountain Science Shinshu University Matsumoto Japan