1. School of Ecology and Nature Conservation Beijing Forestry University Beijing China
2. Institute of Forestry and Climate Change Research Beijing Forestry University Beijing China
3. Sichuan Wolong Forest Ecosystem Research Station Sichuan Academy of Forestry Chengdu China
4. Ecological Restoration and Conservation on Forest and Wetland Key Laboratory of Sichuan Province Sichuan Academy of Forestry Chengdu China
5. College of Agricultural and Life Sciences Kunming University Kunming China