1. MARE—Marine and Environmental Sciences Centre, ESTMInstituto Politécnico de Leiria, 2520‐641 Peniche Portugal
2. Instituto Superior de Agronomia, LEAF, Linking Landscape, Environment, Agriculture and FoodUniversidade de Lisboa, Tapada da Ajuda, 1349‐017 Lisbon Portugal
3. Montpellier SupAgro, UMR 1208 IATE, 2 Place Viala, F‐34060 Montpellier cedex France