1. ARC Centre of Excellence for Coral Reef Studies; James Cook University; Townsville Qld Australia
2. Lancaster Environment Centre; Lancaster University; Lancaster UK
3. Australian Institute of Marine Science; Townsville Qld Australia
4. Department of Mathematics and Statistics; Dalhousie University; Halifax NS Canada
5. College of Marine and Environmental Sciences; James Cook University; Townsville Qld Australia
6. CRIOBE-USR 3278; CNRS-EPHE-UPVD and Laboratoire d'Excellence “CORAIL”; Perpignan Cedex France