1. Key Laboratory of Southwest China Wildlife Resources Conservation (China West Normal University); Ministry of Education; Nanchong Sichuan China
2. Guangxi Institute of Botany; Chinese Academy of Sciences; Guilin Guangxi China
3. Centre of Excellence for Soil Biology; College of Resources and Environment; Southwest University; Beibei Chongqing China
4. School of Plant Biology; University of Western Australia; Crawley WA Australia