1. Institut Botànic de Barcelona (CSIC-ICUB), Passeig del Migdia, s.n,; Catalonia 08038 Barcelona Spain
2. Laboratory of Flora, Institute for Biology and Soil, Academy of Sciences, Chui Str. 265,; Bishkek 720071 Kyrgyzstan
3. Department of Environmental and Plant Biology, ; Plant Development and Evolution, Ohio University,; Porter Hall 500, Athens Ohio 45701 U.S.A.
4. Both authors contributed equally to the design and supervision of this work.