1. Department of Microbiology; School of Preclinical Medicine; Guangxi Medical University; Nanning Guangxi China
2. Guangxi Zhuang Autonomous Region Center for Disease Prevention and Control; Nanning Guangxi China
3. Guangxi Key Laboratory for the Prevention and Control of Viral Hepatitis; Nanning Guangxi China
4. Division of Medicine; UCL Medical School; London UK
5. Department of Public Health of Guangxi Zhuang Autonomous Region; Nanning Guangxi China