1. CESAM and Department of EnvironmentUniversity of Aveiro Aveiro 3810‐193 Portugal
2. CESAM and Department of BiologyUniversity of Aveiro Aveiro 3810‐193 Portugal
3. IPMA Division of Environmental Oceanography and Bioprospecting, Lisbon 1449‐006 Portugal
4. CIIMARUniversity of Porto Matosinhos 4450‐208 Portugal