1. Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden, Chinese Academy of Sciences Wuhan P. R. China
2. Center for Plant Ecology, Core Botanical Gardens Chinese Academy of Sciences Wuhan P.R. China
3. School of Geographic and Tourism Jiaying University Meizhou China
4. Guangdong Provincial Key Laboratory of Conservation and Precision Utilization of Characteristic Agricultural Resources in Mountainous Areas Meizhou P. R. China