1. Shaanxi Key Laboratory for Animal Conservation Northwest University Xi'an China
2. College of Life Science Northwest University Xi'an China
3. Institute of Population and Health Northwest University Xi'an China
4. School of Human Sciences, Centre for Evolutionary Biology The University of Western Australia Perth WA Australia
5. School of Biological Sciences The University of Western Australia Perth WA Australia
6. Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education Northwest University Xi'an China