1. National Institute of Mathematical and Biology Synthesis University of Tennessee Knoxville Tennessee 37996 USA
2. Center for Modeling Complex Interactions University of Idaho 875 Perimeter Drive Moscow Idaho 83844 USA
3. Department of Wildlife Ecology and Conservation University of Florida Gainesville Florida 32611 USA
4. Environmental Sciences Division Oak Ridge National Laboratory P.O. Box 2008 Oak Ridge Tennessee 37830 USA