1. Research Centre in Evolutionary Anthropology and Palaeoecology, School of Natural Sciences and Psychology, Faculty of Science; Liverpool John Moores University; Liverpool L3 3AF United Kingdom
2. Centre for Anatomical and Human Sciences, Hull York Medical School; University of York; York YO10 5DD United Kingdom
3. Department of Earth Sciences; The Natural History Museum; London SW7 5BD United Kingdom
4. Dipartimento di Biologia Ambientale; Sapienza Università di Roma; Roma 00185 Italy