1. Department of Biomedical Sciences A. A. Dugoni School of Dentistry, University of the Pacific San Francisco California USA
2. Department of Biology Saint Mary's College of California Moraga California USA
3. Institut de Paléontologie Humaine, Fondation Albert Ier Prince de Monaco Paris France
4. CEPAM ‐ UMR 7264 CNRS, Université de Nice Nice Cedex 4 France