1. Osaka University; Department of Bioinformatic Engineering, Graduate School of Information Science and Technology; 1-5 Yamadaoka Suita Osaka 565-0871 Japan
2. RIKEN; Quantitative Biology Center; 6-2-3 Furuedai Suita Osaka 565-0874 Japan
3. Tokyo Institute of Technology; Department of Bioengineering; 4259 Nagatsuta-cho Midori-ku, Yokohama 226-8501 Japan
4. RIKEN; Center for Sustainable Resource Science; 1-7-22 Suehiro Tsurumi, Yokohama Kanagawa 230-0045 Japan