1. NC Cooperative Fish and Wildlife Research Unit; Department of Applied Ecology; North Carolina State University; Raleigh NC USA
2. School of Environmental and Forest Science; University of Washington; Seattle WA USA
3. U.S. Geological Survey; NC Cooperative Fish and Wildlife Research Unit; Department of Applied Ecology; North Carolina State University; Raleigh NC USA
4. U.S. Geological Survey; Patuxent Wildlife Research Center; Laurel MD USA