1. Department of Neurology, Klinikum rechts der IsarTechnical University of MunichMunich Germany
2. Institute of Neuronal Cell BiologyTechnical University of MunichMunich Germany
3. Munich Cluster for Systems Neurology (SyNergy)Munich Germany
4. Department of Neurology, Division of Clinical and Experimental NeuroimmunologyUniversity of PecsPecs Hungary
5. Department of Neurology and Institute of Clinical ResearchOdense University Hospital, University of Southern DenmarkOdense Denmark
6. Department of Pathology and ImmunologyUniversity of GenevaGeneva Switzerland
7. Division of Clinical PathologyGeneva University HospitalGeneva Switzerland
8. Department of NeuropathologyUniversity Medical CenterGöttingen Germany
9. Departments of Neurology and Ophthalmology, Program in NeuroscienceUniversity of Colorado Denver School of MedicineAurora CO
10. German Center for Neurodegenerative Diseases (DZNE)Munich Germany
11. Center of Integrated Protein Science (CIPSM)Munich Germany
12. equal contributing senior authors