1. Graduate School of Frontier Sciences; The University of Tokyo; 5-1-5 Kashiwanoha, Kashiwa Chiba 277-8561 Japan
2. Center for Environmental Risk Research; National Institute for Environmental Studies; 16-2 Onogawa, Tsukuba Ibaraki 305-8506 Japan
3. Okazaki Institute for Integrative Bioscience, National Institute for Basic Biology, and Department of Basic Biology; School of Life Science, Graduate University for Advanced Studies (SIKENDAI); 5-1 Higashiyama, Myodaiji Okazaki Aichi 444-8787 Japan