1. Restoration and Conservation Biology Research Group, School of Animal, Plant and Environmental SciencesUniversity of the Witwatersrand Johannesburg, Private Bag 3 WITS 2050 Johannesburg South Africa
2. Centre for African Ecology, School of Animal, Plant and Environmental SciencesUniversity of the Witwatersrand Johannesburg, Private Bag 3 WITS 2050 Johannesburg South Africa
3. School of Forestry, Environmental and Geographical SciencesMakerere University PO Box 7062 Kampala Uganda