1. Department of Genetics; Geisel School of Medicine; Dartmouth College; Hanover New Hampshire United States of America
2. Institute of Quantitative Biomedical Sciences; Dartmouth College; Hanover New Hampshire United States of America
3. Division of Computational Bioscience; Center for Information Technology; National Institutes of Health; Bethesda Maryland United States of America
4. Department of Community and Family Medicine; Geisel School of Medicine; Dartmouth College; Hanover New Hampshire United States of America