1. Department of NeurosciencePerelman School of Medicine at the University of PennsylvaniaPhiladelphia PA
2. Department of Clinical Studies–New Bolton CenterUniversity of Pennsylvania School of Veterinary MedicinePhiladelphia PA
3. Department of NeurologyPerelman School of Medicine at the University of PennsylvaniaPhiladelphia PA
4. August Pi i Sunyer Institute of Biomedical Research and Catalan Institute for Research and Advanced Studies, Hospital Clinic, University of BarcelonaBarcelona Spain