1. Department of Horticultural Science; Faculty of Agriculture; Kagoshima University; Kagoshima Japan
2. Kagoshima University Experimental Farm; Kagoshima Japan
3. Department of Bioscience and Biotechnology; Faculty of Agriculture; Kyushu University; Fukuoka Japan
4. Faculty of Education and Integrated Arts and Sciences; Waseda University; Tokyo Japan
5. Department of Biology; Faculty of Science; Kyushu University; Fukuoka Japan