1. School of Anatomical Sciences, Faculty of Health Sciences University of the Witwatersrand Johannesburg Republic of South Africa
2. Center for Zoo and Wild Animal Health, Copenhagen Zoo Frederiksberg Denmark
3. Borås Zoo Västergötland Borås Sweden
4. Nash Family Department of Neuroscience and Friedman Brain Institute Icahn School of Medicine at Mount Sinai New York New York USA
5. New York Consortium in Evolutionary Primatology New York New York USA
6. Department of Anatomy Des Moines University Des Moines Iowa USA