1. Key Lab of Chinese Medicine Resources Conservation, State Administration of Traditional Chinese Medicine of the People's Republic of China, Institute of Medicinal Plant Development; Chinese Academy of Medical Sciences, Peking Union Medical College; Beijing China
2. Chongqing Institute of Medicinal Plant Cultivation, Research and Utilization on Characteristic Biological Resources of Sichuan and Chongqing Co-construction Lab; Chinese Medicine Breeding and Evaluation Engineering Technology Research Center of Chongqing; Chongqing China
3. Beijing Botanical Garden, Institute of Botany; Chinese Academy of Sciences; Beijing China