1. State Key Laboratory of Vegetation and Environmental Change; Institute of Botany, the Chinese Academy of Sciences; 20 Nanxincun, Xiangshan Beijing 100093 PR China
2. Beijing Museum of Natural History; Beijing 100050 PR China
3. Chinese Research Academy of Environmental Sciences; 8 Dayangfang Beijing 100012 PR China
4. College of Life Science; Linyi University; Linyi Shandong Province 276005 PR China