1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing China
2. CAS Center for Excellence in Life and Paleoenvironment Beijing China
3. Division of Paleontology American Museum of Natural History New York NY USA
4. Earth and Environmental Sciences Graduate Center City University of New York New York NY USA
5. School of Earth Sciences University of Bristol Bristol UK
6. Department of Ecology and Evolutionary Biology Brown University Providence RI USA
7. College of Earth and Planetary Sciences University of Chinese Academy of Sciences Beijing China