1. Dental Physical Sciences, Barts and The London School of Medicine and Dentistry, Queen Mary University of LondonLondonE1 4NS United Kingdom
2. Oxford Animal Flight Group, Department of Zoology, University of OxfordOxfordOX1 3PS United Kingdom
3. Structure and Motion LaboratoryDepartment of Comparative Biomedical SciencesThe Royal Veterinary CollegeHatfield HertfordshireAL9 7TA United Kingdom
4. Skeletal BiologyDepartment of Comparative Biomedical SciencesThe Royal Veterinary CollegeLondonNW1 0TU United Kingdom