1. Department of Zoology and Entomology Rhodes University Grahamstown South Africa
2. Department of Zoology Nelson Mandela University Gqeberha South Africa
3. LOG Laboratoire d'Océanologie et de Géosciences CNRS, UMR 8187 Univ. LilleUniv. Littoral Côte d'Opale Wimereux France
4. Department of Marine Resources and Energy Tokyo University of Marine Science and Technology Tokyo Japan