1. Department of Biology University of Toronto at Mississauga Mississauga Ontario Canada
2. Station d'Ecologie Théorique et Expérimentale, UAR 2029, CNRS Moulis France
3. Environment and Sustainability Institute University of Exeter Penryn
4. School of Biological Sciences University of Reading Reading UK
5. Institute of Marine Ecosystem and Fishery Science, Centre for Earth System Research and Sustainability Universität Hamburg Hamburg Germany