1. Department of Environmental Sciences, Zoological Institute University of Basel Basel 4051 Switzerland
2. School of Biology, Centre for Biological Diversity University of St Andrews St Andrews KY16 9TH United Kingdom
3. Natural History Museum University of Oslo Oslo 0318 Norway
4. CNRS, ECOBIO (Écosystèmes, Biodiversité, Évolution) – UMR 6553 Université de Rennes 1 Rennes 35000 France
5. Evolutionary Biology Bielefeld University Bielefeld DE‐33615 Germany