1. Program in Ecology, Evolution, and Conservation Biology; University of Nevada; Reno/MS 436 Reno NV 89557 USA
2. Owyhee Desert Studies; 18109 Briar Creek Road Murphy ID 83650 USA
3. U.S. Geological Survey; Forest and Rangeland Ecosystem Science Center; Snake River Field Station Boise ID 83706 USA
4. Department of Biology; Boise State University; 1910 University Drive Boise ID 83725 USA