1. Institute of Translational and Clinical Medicine, Medical School Newcastle University Newcastle upon Tyne UK
2. Medicines Research Centre GlaxoSmithKline London UK
3. Biosciences Institute, Medical School Newcastle University Newcastle upon Tyne UK
4. Department of Biosciences Durham University Durham UK
5. Department of Dermatology, Royal Victoria Infirmary and NIHR Newcastle Biomedical Research Centre Newcastle Hospitals NHS Foundation Trust Newcastle upon Tyne UK