1. University of Lyon, University Claude Bernard Lyon 1; UMR-CNRS 5023 LEHNA; 43 Boulevard du 11 Novembre 1918 69622 Villeurbanne Cedex France
2. Department of Life, Health and Environmental Sciences; University of L'Aquila; Via Vetoio, Coppito I-67100 L'Aquila Italy
3. IMBE, Institut Méditerranéen de Biodiversité et Ecologie marine et continentale; UMR CNRS 7263-Aix Marseille University, Avignon University, CNRS, IRD, IMBE; Marseille France
4. Invertebrate Section; Royal Belgian Institute of Natural Sciences; Vautierstraat, 29 B-1000 Brussels Belgium