1. Department of Biology, Ecology and Evolution University of Fribourg Chemin du Musée 10 1700 Fribourg Switzerland
2. Tvärminne Zoological Station J.A. Palménin tie 260 10900 Hanko Finland
3. Department of Genetics, Evolution and Environment University College LondonInstitute of Healthy Ageing Darwin Building, Gower Street London WC1E 6BT United Kingdom
4. CEFE, Univ Montpellier, CNRS, EPHE, IRD, Univ Paul Valéry Montpellier 3 1919, route de Mende 34293 Montpellier Cedex 5 France