1. Institute of Livestock and Grassland Science National Agriculture and Food Research Organization (NARO) Tsukuba Ibaraki Japan
2. Forestry and Forest Products Research Institute Tsukuba Ibaraki Japan
3. Center for Environmental Biology and Ecosystem Studies National Institute for Environmental Studies Tsukuba Ibaraki Japan
4. School of Veterinary Medicine Azabu University Sagamihara Kanagawa Japan
5. Insutitute for Biodiversity Research and Education Earthworm Karuizawa Nagano Japan
6. Advanced Analysis Center NARO Tsukuba Ibaraki Japan
7. Faculty of Science and Engineering Chuo University Bunkyo‐ku Tokyo Japan